{# load django_bootstrap5#%} CCDB server
Prof. Itay Mayrose Lab - Plant Evolution, bioinformatics, & comparative genomics


   CCDB is a community resource. If you are aware of any missing data or if any of the counts are erroneous please submit it to the forms submit new counts, or report erroneous counts.


CCDBPteridophytesCystopteridaceaeCystopterisCystopteris fragilis

120 chromosome counts in Cystopteris fragilis:


Name Accepted Name Gametophytic(n) Sporophytic(2n) Data Source reference
  Cystopteris altajensis Gureeva Cystopteris fragilis var. altajensis (Gureeva) Stepanov 126   IAPT/IOPB IAPT/IOPB Chromosome Data 26
!   Cystopteris diaphana (Bory) Blasdell Cystopteris fragilis (L.) Bernh.   252 Love, 1977 Blasdell 1963
!   Cystopteris diaphana (Bory) Blasdell Cystopteris fragilis (L.) Bernh.   252 Love, 1977 Manton, Lovis & Vida (in Vida 1974)
  Cystopteris diaphana (Bory) Blasdell Cystopteris fragilis (L.) Bernh. 42, 126   Cave1963 Blasdell 1963
!   Cystopteris dickieana R. Sim Cystopteris fragilis (L.) Bernh.   n=84 British Isles cytology Manton, I. Probl. Cytol. Evol. Pterid. (1950)
!   Cystopteris dickieana R. Sim Cystopteris fragilis (L.) Bernh.   168 flora of NW Europe http://wbd.etibioinformatics.nl/bis/flora.php?selected=beschrijving&menuentry=soorten&id=1517
!   Cystopteris dickieana R. Sim Cystopteris fragilis (L.) Bernh.   168 Flora Europaea Manton, 1950
!   Cystopteris dickieana R. Sim Cystopteris fragilis (L.) Bernh. 126   IPCN65 Manton & R. 1965
  Cystopteris dickieana R. Sim Cystopteris fragilis (L.) Bernh. 84   IPCN online Rasbach, H. 1998. In R. Wisskirchen & H. Haeupler, Standardliste der Farn- und Blütenpflanzen Deutschlands. Bundesamt für Naturschutz & Verlag Eugen Ulmer, Stuttgart.
  Cystopteris dickieana R. Sim Cystopteris fragilis (L.) Bernh.   168 IPCN online Obermayer, R., I. J. Leitch, L. Hanson & M. D. Bennett. 2002. Nuclear DNA C-values in 30 species double the familial representation in pteridophytes. Ann. Bot. (Oxford) 90: 209–217.
!   Cystopteris dickieana R. Sim Cystopteris fragilis (L.) Bernh. 126   IPCN67-71 BIR, s.s., 1971. Cytological observations on some Indian species of Cystopteris Bernh. Nucleus 14: 56-62.
!   Cystopteris dickieana R. Sim Cystopteris fragilis (L.) Bernh.   168 IPCN75-78 Love & Love 1976
!   Cystopteris dickieana R. Sim Cystopteris fragilis (L.) Bernh.   168 Love, 1977 Manton 1950
!   Cystopteris dickieana R. Sim Cystopteris fragilis (L.) Bernh.   168 Love, 1977 Wagner 1955
!   Cystopteris dickieana R. Sim Cystopteris fragilis (L.) Bernh.   168 Love, 1977 Vida 1974
!   Cystopteris dickieana R. Sim Cystopteris fragilis (L.) Bernh.   168 Love, 1977 Love & Love (in Love 1976)
!   Cystopteris dickieana R. Sim Cystopteris fragilis (L.) Bernh.   168,252 eFlora  
!   Cystopteris dickieana R. Sim Cystopteris fragilis (L.) Bernh. 84   Cave1956supp Wagner 1955
!   Cystopteris fragilis Cystopteris fragilis (L.) Bernh. 84   CromoCat 2015 Queirós, M. (1991b). Estudos cromossómicos em Pteridophyta de Portugal. II. Collect. Bot. (Barcelona) 20: 23-33.
!   Cystopteris fragilis Cystopteris fragilis (L.) Bernh.   168 CromoCat 2015 Queirós, M. (1991b). Estudos cromossómicos em Pteridophyta de Portugal. II. Collect. Bot. (Barcelona) 20: 23-33.
!   Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   n=84 British Isles cytology Manton, I. Probl. Cytol. Evol. Pterid. (1950)
!   Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   n=126 British Isles cytology Manton, I. Probl. Cytol. Evol. Pterid. (1950)
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   168 Flora Iberica Cytotaxonomy of spanish plants. I. Introduction. Pteridophyta and Gymnospermae. Lve, A. & E. Kjellqvist.Lagascalia2(1): 23-35(1972).
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh. 84   Flora Iberica Estudos cromossmicos em Pteridophyta de Portugal. II. Queirs, M.Collect. Bot. (Barcelona)20: 23-33(1991).
!   Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   168, 252 flora of NW Europe http://wbd.etibioinformatics.nl/bis/flora.php?selected=beschrijving&menuentry=soorten&id=1516
!   Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   168 Flora Europaea Manton, 1950
!   Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   252 Flora Europaea Manton, 1950
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh. c.84   IPCN65 Walker 1964-65
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh. 84   IPCN online Haufler, C. H. & D. E. Soltis. 1986. Genetic evidence suggests that homosporous ferns with high chromosome numbers are diploid. Proc. Natl. Acad. Sci. U.S.A. 83: 4389–4393.
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh. 84   IPCN online Kurita, S. 1986. Chromosome studies on South American pteridophytes (1). In: Contributions to the Botany of the Andes. (II) 47–67.
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh. 42II   IPCN online Tsai, J. L. & W. C. Shieh. 1985. A cytotaxonomic survey of the fern family Aspidiaceae (sensu Copeland) in Taiwan. J. Sci. Engin. 22: 121–144.
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh. 84   IPCN online Dalgaard, V. 1989. Additional chromosome numbers in vascular plants from the Disko Bugt area (west Greenland). Willdenowia 19: 199–213.
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   168 IPCN online Love, A. & D. Love. 1976. In IOPB chromosome number reports LIII. Taxon 25: 483–500.
!   Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   c.168 IPCN online Obermayer, R., I. J. Leitch, L. Hanson & M. D. Bennett. 2002. Nuclear DNA C-values in 30 species double the familial representation in pteridophytes. Ann. Bot. (Oxford) 90: 209–217.
!   Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh. 53II+60I+2III   IPCN online Paler, M. H. & D.S. Barrington. 1995. The hybrid Cystopteris fragilis x C. tenuis (Dryopteridaceae) and the relationship between its tetraploid progenitors. Syst. Bot. 20: 528–545.
!   Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh. 80-85   IPCN66 Taylor & Brockman 1966
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   168 IPCN72 LOEVE, A., & E. KJELLQVIST. 1972. Cytotaxonomy of Spanish plants. I. Introduction. Pteridophyta and Gymnospermae. Lagascalia 2: 23-35.
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh. 84   IPCN67-71 TAYLOR, R.L., & G.A. MULLIGAN. 1968. Flora of the Queen Charlotte Islands. Part 2. Cytological aspects of the Vascular Plants. Queen's Printer, Ottawa. pp. 148.
!   Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh. 126   IPCN67-71 70-309
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh. 84   IPCN67-71 BIR, S.S. & P. SHUKLA. 1967. Cytology of some north Indian ferns. Cytologia 32: 24-30.
!   Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   168 IPCN75-78 Love & Love 1976
!   Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   168 Love, 1977 Manton 1950
!   Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   168 Love, 1977 Britton 1953
!   Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   168 Love, 1977 Wagner 1955
!   Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   168 Love, 1977 Bir (in Mehra 1961b)
!   Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   168 Love, 1977 Love & Love 1961a
!   Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   168 Love, 1977 Sorsa 1961
!   Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   168 Love, 1977 Sorsa 1962
!   Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   168 Love, 1977 Blasdell 1963
!   Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   168 Love, 1977 Britton (in Fabbri 1963)
!   Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   168 Love, 1977 Wagner (in Fabbri 1963)
!   Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   168 Love, 1977 Britton 1964
!   Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   c.168 Love, 1977 Taylor, R. L. & Brockman 1966
!   Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   168 Love, 1977 Bir (in Love 19670)
!   Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   168 Love, 1977 Bir & Shukla 1967
!   Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   168 Love, 1977 Taylor, R. L. & Mulligan 1968
!   Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   168 Love, 1977 Bir 1971b
!   Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   168 Love, 1977 Love & Kjellvist 1972
!   Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   168 Love, 1977 Smith, A. R. 1974
!   Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   168 Love, 1977 Vida 1974
!   Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   168 Love, 1977 Love & Love (in Love 1976)
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   168 IPCN73-74 SMITH, A.R. 1974. Taxonomic and cytological notes on ferns from California and Arizona. Madrono 22: 376-378.
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh. 76 II + 581   IPCN73-74 VIDA, G. 1974. Genome analysis of the European Cystopteris fragilis complex. I. Tetraploid taxa. Acta Bot. Acad. Sci. Hung. 20: 181 192
!   Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh. c.84 II + c.42 I   IPCN73-74 VIDA, G. 1974. Genome analysis of the European Cystopteris fragilis complex. I. Tetraploid taxa. Acta Bot. Acad. Sci. Hung. 20: 181 192
!   Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   168,252,336,84 eFlora  
!   Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh. n = 126   Austria_checklist -- 1974. Genome analysis of the European Cystopteris fragilis complex. I. Tetraploid taxa. Acta Bot.Acad.Sci.Hung 20[1-2], 181-192.°
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh. 84   Cave1956supp Wagner 1955
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh. 84 ± 2   Cave1956supp Britton 1953
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh. 84   Cave1962 Sorsa 1961
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh. 84   Cave1963 Britton in Fabbri 1963; Wagnei in Fabbri 1963; Sorsa 1962
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh. 84, c.126, 126 126 Cave1963 Blasdell 1963
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh. c.130   CromoCat 2015 Favarger, C., Galland, N. & Küpfer, P. (1979). Recherches cytotaxonomiques sur la flore orophile du Maroc. Naturalia Monspel., Sér. Bot. 29: 1-64.
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh. c.126   CromoCat 2015 Blasdell, R. F. (1963). A monographic study of the fern genus Cystopteris. Mem. Torrey Bot. Club 21: 1-102.
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh. 84   CromoCat 2015 Bir, S. S. & Shukla, P. (1967). Cytology of some north Indian ferns. Cytologia 32: 24-30.
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh. 84   CromoCat 2015 Bir, S. S. (1971). Cytological observations on some Indian species of Cystopteris Bern. Nucleus 14: 56-62.
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh. 84   CromoCat 2015 Blasdell, R. F. (1963). A monographic study of the fern genus Cystopteris. Mem. Torrey Bot. Club 21: 1-102.
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh. 84   CromoCat 2015 Löve, A. & Löve, D. (1961b). Some chromosome numbers of Icelandic ferns and fern-allies. Amer. Fern. J. 51: 127-128.
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh. 84   CromoCat 2015 Manton, I. (1950). Problems of cytology and evolution in the Pteridophyta. Cambridge
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh. 84   CromoCat 2015 Mehra, P. N. & S. P. Khullar. (1974). Cytotaxomomy of W. Himalayan ferns. II. Bull. Panjab Univ. 25: 135-178.
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh. 84   CromoCat 2015 Smith, A. R. & J. T. Mickel. (1977). Chromosome counts for Mexican ferns. Brittonia 29: 391-398.
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh. 84   CromoCat 2015 Sorsa, V. (1961). Chromosome studies in Finnish Pteridophyta, II. Hereditas 47: 480-488.
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh. 84   CromoCat 2015 Sorsa, V. (1962). Chromosomenzahlen Finnischer Kormophyten, I. Ann. Acad. Sci. Fenn., Sér. A 58: 1-14.
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh. 84   CromoCat 2015 Taylor, R. L. & G. A. Mulligan. (1968). Flora of the Queen Charlotte Island. Part 2. Cytologiacl aspects of the Vascular Plants. Queen's Printer Ottawa
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh. 42II   CromoCat 2015 Tsai, J. L. & W. C. Shieh (1985). A cytotaxonomic survey of the fern family Aspidiaceae (sensu Copeland) in Taiwan. J. Sci. Engin. (Taiwan) 22: 121-144
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh. 126   CromoCat 2015 Mitui, K. (1970). Chromosome studies on Japanese ferns (4). J. Jap. Bot. 45: 84-90.
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   168 CromoCat 2015 Britton, D. M. (1953). Chromosome studies on ferns. Amer. J. Bot. 40: 575-583.
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   168 CromoCat 2015 Brownlie, G. (1958). Chromosome numbers in New Zealand ferns. Trans. Roy. Soc. New Zealand 85: 213-216.
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   168 CromoCat 2015 Löve, A. & Kjellqvist, E. (1972). Cytotaxonomy of spanish plants. I. Introduction. Pteridophyta and Gymnospermae. Lagascalia 2(1): 23-35.
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   168 CromoCat 2015 Löve, A. & Löve, D. (1976). Reports [In Löve, A. (ed.), IOPB chromosome numbers reports LIII]. Taxon 25(4): 484-487.
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   168 CromoCat 2015 Vida, G. (1974). Genome analysis of the European Cystopteris fragilis complex. I. Tetraploid taxa. Acta Bot. Acad. Sci. Hung. 20: 181-192.
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   168 CromoCat 2015 Wagner, W. H. & D. J. Hagenah. (1956). A diploid variety in the Cystopteris fragilis complex. Rhodora 58: 79-87.
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   168 CromoCat 2015 Wagner, W. H. (1955). Cytotaxonomic observations on North American ferns. Rhodora 57: 219-240.
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   252 CromoCat 2015 Vida, G. (1974). Genome analysis of the European Cystopteris fragilis complex. I. Tetraploid taxa. Acta Bot. Acad. Sci. Hung. 20: 181-192.
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   252 CromoCat 2015 Wagner, W. H. & D. J. Hagenah. (1956). A diploid variety in the Cystopteris fragilis complex. Rhodora 58: 79-87.
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   252 CromoCat 2015 Wagner, W. H. (1955). Cytotaxonomic observations on North American ferns. Rhodora 57: 219-240.
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   c.168 CromoCat 2015 Obermayer, R. & Leitch, I.J. & Hanson, L. & Bennett, M.D. (2002). Nuclear DNA C-values in 30 species double the familial representation in pteridophytes. Annals of Botany 90:209-217.
!   Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh. 53II+60I+2III   CromoCat 2015 Paler, M. H. & D.S. Barrington (1995). The hybrid Cystopteris fragilis x C. tenuis (Dryopteridaceae) and the relationship between its tetraploid progenitors Syst. Bot. 20: 528–545
!   Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh. 126   CromoCat 2015 Castroviejo & al. (2003). Flora Iberica (vol. I-VIII, X, XIV) Publicaciones del CSIC Madrid
!   Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   252 CromoCat 2015 Castroviejo & al. (2003). Flora Iberica (vol. I-VIII, X, XIV) Publicaciones del CSIC Madrid
!   Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh.   336 CromoCat 2015 Castroviejo & al. (2003). Flora Iberica (vol. I-VIII, X, XIV) Publicaciones del CSIC Madrid
  Cystopteris fragilis (L.) Bernh. Cystopteris fragilis (L.) Bernh. 126   IAPT/IOPB IAPT/IOPB Chromosome Data 26
!   Cystopteris fragilis None Cystopteris fragilis (L.) Bernh. 84   Cave1958 Brownlie 1958
  Cystopteris fragilis f. fragilis Cystopteris fragilis (L.) Bernh. 84   IPCN online Bir, S. S., C. K. Satija, P. Goyal & S. M. Vasudeva. 1979--1980. Further studies on the distribution and taxonomy of Cystopteris fragilis (L.) Bernh. from the western Himalayas. Phyta, J. Soc. Pl. Taxomomists 2–3: 62–66.
!   Cystopteris fragilis f. fragilis Cystopteris fragilis (L.) Bernh. 84   IPCN67-71 BIR, s.s., 1971. Cytological observations on some Indian species of Cystopteris Bernh. Nucleus 14: 56-62.
!   Cystopteris fragilis f. fragilis Cystopteris fragilis (L.) Bernh. 84   IPCN67-71 BIR, s.s., 1971. Cytological observations on some Indian species of Cystopteris Bernh. Nucleus 14: 56-62.
!   Cystopteris fragilis f. fragilis Cystopteris fragilis (L.) Bernh. 84   IPCN67-71 BIR, s.s., 1971. Cytological observations on some Indian species of Cystopteris Bernh. Nucleus 14: 56-62.
!   Cystopteris fragilis f. fragilis Cystopteris fragilis (L.) Bernh. 84   IPCN67-71 BIR, s.s., 1971. Cytological observations on some Indian species of Cystopteris Bernh. Nucleus 14: 56-62.
!   Cystopteris fragilis f. simulans Weath. Cystopteris utahensis Windham & Haufler 84   Cave1956supp Wagner 1955
  Cystopteris fragilis subsp. fragilis Cystopteris fragilis (L.) Bernh. 84   IPCN online Queirós, M. 1991. Estudos cromossómicos em Pteridophyta de Portugal. II. Collect. Bot. (Barcelona) 20: 23–33.
  Cystopteris fragilis subsp. fragilis Cystopteris fragilis (L.) Bernh. 84   IPCN online Queirós, M. 1998. Estudos cromosómicos em Pteridophyta de Portugal, III. Collect. Bot. (Barcelona) 23: 73–77.
!   Cystopteris fragilis subsp. fragilis Cystopteris fragilis (L.) Bernh. n = 126   Austria_checklist Blasdell, R. F. 1963. A monographic study of the fern genus Cystopteris. Mem.Torrey Bot.Club 21[4], 1-102. under Cystopteris fragilis (L.) Bernh.
!   Cystopteris fragilis var. dickieana (R. Sim) T. Moore Cystopteris fragilis (L.) Bernh. 126   IPCN online Manton, I., J. D. Lovis, G. Vida & M. GIBBY. 1986. Cytology of the fern flora of Madeira. Bull. Brit. Mus. (Nat. Hist.), Bot. 15(2): 123–161.
!   Cystopteris fragilis var. mackayii G. Lawson Cystopteris tenuis (Michx.) Desv. 84   Cave1956 Wagner & Hagenah
!   Cystopteris fragilis var. protrusa Weath. Cystopteris protrusa (Weath.) Blasdell 42   Cave1956 Wagner & Hagenah
!   Cystopteris fragilis var. tennesseensis McGregor Cystopteris utahensis Windham & Haufler 126   Cave1956supp Wagner 1955
  Cystopteris sikkimensis Ching ex Bir Cystopteris fragilis (L.) Bernh. 168   IPCN67-71 BIR, s.s., 1971. Cytological observations on some Indian species of Cystopteris Bernh. Nucleus 14: 56-62.
!   Cystopteris sikkimensis Ching ex Bir Cystopteris fragilis (L.) Bernh.   336 Love, 1977 Bir (in Mehra 1961b)
!   Cystopteris sikkimensis Ching ex Bir Cystopteris fragilis (L.) Bernh.   336 Love, 1977 Bir 1971b
  Cystopteris viridula (Desv.) Desv. Cystopteris fragilis (L.) Bernh. 84   Flora Iberica Notas cariolgicas e fitogeograficas de algumas Pteridophyta de Portugal. I Queirs, M., J. Ormonde & I. Nogueira.Acta Bot. Malacitana13: 121-140(1988).
  Cystopteris viridula (Desv.) Desv. Cystopteris fragilis (L.) Bernh. 84   IPCN online Queiros, M., J. Ormonde & I. Nogueira. 1988. Notas cariológicas e fitogeograficas de algunas Pteridophyta de Portugal I. Acta Bot. Malac. 13: 121–140.